Using gland regions on their heads, warthogs mark poles and other objects for self-orientation. Males often mark during the mating season. The male also sprays urine when searching for and inspecting sleeping cavities.
1974
2020
1976
2023
—
1959
2017
2010
1967
2003
2015
1962
1961
2004
1934
2021
2024
1895
1980