Using gland regions on their heads, warthogs mark poles and other objects for self-orientation. Males often mark during the mating season. The male also sprays urine when searching for and inspecting sleeping cavities.
1933
1953
1999
1968
1980
1961
2015
—
2004
2014
1895
1896
1959
2002
2016
2013
1944
1997
1996
2022