Using gland regions on their heads, warthogs mark poles and other objects for self-orientation. Males often mark during the mating season. The male also sprays urine when searching for and inspecting sleeping cavities.
2000
2001
2016
2013
2008
2017
1895
2012
1959
2015
1953
1961
1896
—
1968
2002
1999
2018