After uniting the entire Brodenthood under his rule, valiant Emperodent Julius Cheezer sets out to prove his worth by stealing a forbidden block of cheese from the dreaded Realm of Giants - defying the pleas of his squeakheart Cleoratra. His curdsade takes an unexpected turn when he encounters Romeo Montague, a desolate human coming off a certain life tragedy. Together, they explore the meaning of love, legacy, and life itself.