Fahri Fırfır, a hotel owner blinded by greed, and his manager Şehnaz Şıkır want to expand their hotels by destroying the forests. A family defying them finds an unexpected ally: the mole team consisting of Boyo, Süslü, Kösteban, Köstepem, and Köstosun will do everything in their power to protect their forest.

Pelin and Caner are the closest friends of the Mole family. Shadow, who was turned into a tree because of the evil deeds he had done in the past, was saved by Uncle Köstan of the Mole family in exchange for his promise to become a good person. However, Shadow broke his promise and stole Queen Sora's dress, which was said to be enchanted, from the auction house where he was working. As soon as he touched the dress, the evil inside it activated the tree charm that Uncle Köstan had previously made. Gölge began to turn back into a tree. He was found in the forest by Şahane Postum, a hunter who was fond of precious stones and furs. Shahane is immediately captivated by Queen Sora's dress and desires to possess it. However, the enchantment from Uncle Köstan, which has transferred into the dress, begins to transform Shahane into a deer as well. The Shadow suggests that they must go to Uncle Köstan to resolve the situation. However, Uncle Köstan has already set off toward Köstebekya.